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Butterflies

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NABOKOV'S SATYR

Nabokov’s Satyr, Ramsey Canyon, AZ (Mike Watson)

Researching range-restricted, spring-flying butterflies that I might see on my recent California & Arizona Specialities Birdquest tour I was struck by the fact that most of the special butterflies fly in the Arizona ‘second spring’ monsoon season from July to September. However, there are a few that could be seen in the actual springtime of my visit, in early May. One of these caught my eye, Nabokov’s Satyr. Not only as it carries a famous name, but I noted that it belonged to a new butterfly genus for me Cyllopsis, the ‘gemmed satyrs’. Essentially subtle browns, with jewel-like studs embedded on their hindwing borders, like blues. What a wonderful combination! There are more than 30 members of this Neotropical genus, three of which occur in the southern United States: Gemmed Satyr Cyllopsis gemma; Canyonland Satyr C. pertepida and Nabokov’s Satyr C. pyracmon. My pal Craig Robson, who spends part of the year in Southeast Arizona, said the latter was ‘easy anywhere in the mountains on trails/in shade’ and looking at the number of INaturalist.org sightings I thought I should have no trouble in catching up with it in early May. With many pairs of eyes in my group I mentioned that we should be on the lookout for any small brown butterflies and sure enough on our first morning in the Sky Island canyons, at the lovely Madera Canyon near Green Valley, while returning from a successful hike in search of Coppery-tailed Trogon, sharp-eyed Richard spotted one. It was a Nabokov’s, albeit a worn individual with a torn hindwing. Very exciting nevertheless! Thanks Richard! After alighting on a stone, it moved off quickly uphill and into dense scrub and out of reach. Moving with surprising speed at ground level, it skipped through the bases of thorn bushes, a most annoying habit of this species I was to discover in the coming days.

Nabokov’s Satyr, Madera Canyon, AZ (Mike Watson)

Interest in butterflies on our birding tours is very limited, so my opportunities are therefore compressed into a few minutes here and there. The next canyon bottom we visited, the lovely Miller Canyon also produced Nabokov’s Satyr at each stop, but in the heat of the midday they were completely restless and rarely paused for more than a few seconds on their ramblings through the shady understorey of the evergreen oak and Alligator Juniper woodland, when they would angle their wings perpendicular to the shafts of sunlight reaching the leaf litter, in the manner of a grayling back home. This did not allow me enough time to get in position for a photo before they were off again. Goodness me, it was hard work following these pesky creatures round and round a small patch of woodland, before ultimately losing contact, as they made off through the bushes. All too soon, my group returned from a hummingbird feeder sojourn with my co-leaders and that was that.

Nabokov’s Satyr, Ramsey Canyon, AZ (Mike Watson)

They were indeed in each canyon and, finally, it was in Ramsey Canyon, the most famous of all the birding spots in the Huachuca Mountains, that I could finally spend some quality macro time with the satyrs, significantly late in the afternoon when they settled down to visit wet patches of ground in the watercourses and directly under the hummingbird feeders! Most were quite worn by this stage in their flight season and it was more difficult to read their undersides to separate them from the very similar Canyonland Satyr (which is usually found further uphill rather than in the valley bottom) but they were very approachable now. A very satisfying experience to go alongside the other avian treasures of these precious habitats. How many takers for ‘I’ve got a settled Nabokov’s Satyr here’? None at all. We are so few.

Madera Canyon, Arizona - Nabokov’s Satyr habitat, oak and Alligator Juniper woodland (Mike Watson)

The butterfly is named after the Russian/American author Vladimir Nabokov (1899 – 1977), a fascinating character whose most famous work is the 1955 novel Lolita. It is hailed one of the greatest literary works of the 20th century but the subject is such a hot potato now! He was born of Russian nobility but moved gradually westwards, fleeing the Russian Revolution and then Hitler’s Germany, before settling in the United States and gaining citizenship in 1945. Nabokov was a true polymath and as well as an acclaimed author, he was also an accomplished entomologist and has several butterfly and moth species named after both him and his literary works! He wrote in three languages; Russian, English and French, gave boxing lessons and composed chess problems. He lectured at Cornell, (where one of his students was US Supreme Court Justice Ruth Bader Ginsberg). He was mostly concerned with studying blue butterflies and genetic research later supported his hypothesis that the Polyommatus blues came to the Americas over the Bering Strait in five waves, eventually reaching Chile. I was fascinated to read that Nabokov did not learn to drive, he got his wife Véra to drive him around in search of butterflies and in 1967, he commented ‘The pleasures and rewards of literary inspiration are nothing beside the rapture of discovering a new organ under the microscope or an undescribed species on a mountainside in Iran or Peru. It is not improbable that had there been no revolution in Russia, I would have devoted myself entirely to lepidopterology and never written any novels at all.’ Interestingly ‘his’ satyr was first described in 1867 and the use of his name stems from the description of its subspecies named nabokovi in his honour by Lee D. Miller in 1974, from none other than Ramsey Canyon, Arizona (!). The epithet Nabokov’s was later adopted as the common name for the larger species. I hope it does not get cancelled, as is the current Cornell habit with birds. We would surely end up with some hopeless anodyne alternative.

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FALSE COMMA


False Comma Nympahlis vau-album - showing its white ‘V’. One of Europe’s rarest butterflies!

The False Comma Nymphalis vau-album (or Compton Tortoiseshell, as it is known in North America) has always been a scarce butterfly on the western limit of its range in Europe. In recent years it has become even more so, and is considered extinct in Austria, Czech Republic, Estonia, Hungary, Slovenia and Slovakia. However, it seems to be holding its own in the mountains of southeast Serbia and a number of butterfly tour companies now target this spectacular nymphalid. I was privileged to join my friend János Oláh’s 2025 Sakertours Serbia butterfly watching tour. János applies the same level of determination and planning to finding rare butterflies as he does to birds and we had a fantastic time, recording 110 species in five full days including some very sought after ones. As well as False Comma, in this small corner of Serbia, near the Bulgarian border, we also saw: Yellow-banded Skipper Pyrgus sidae; Apollo Parnassius apollo; Violet Copper Lycaenea helle (among seven species of copper!); Blue Argus Aricia anteros; Bog Fritillary Boloria eunomia (among seventeen species of fritillary); Poplar Admiral Limentis populi (four, of the all dark form tremulae, prevalent in the Balkans); the enigmatic Lesser Lattice Brown Kirinia climene; Russian Heath Coenonympha leander; Bulgarian Ringlet Erebia orientalis and Almond-eyed Ringlet Erebia albergana, to mention a few.

This False Comma has a small notch in its left wing and was easily identified as the same individual.

János has done a lot of testing over the last few years, checking the optimum timing for this tour and he concluded that 11 June was a good start date for freshly emerged butterflies of a wide range of species hopefully including False Comma at the beginning of its season. However, after one of the coldest springs on record in eastern Europe this year we were nervous that we might be too early this time. We need not have worried; it seemed that the recent warm weather just before the tour start had got things back on track for some normal flight timings. There are zillions of waypoints available online for previous sightings of False Comma, but it is a mobile butterfly, so it pays to know some of its favourite spots. There is much else to see in this part of Serbia and János promised all the other butterflies as well, so we did not devote 100% of our time to searching for False Comma. There is evidence that it is mostly active in Serbia between 8.30AM and 12PM in warm conditions in June/July, with ‘virtually no activity after midday’ (Gascoigne-Pees et al 2014) and we spent plenty of time in known previous locations around this time. So, we were interested to get a hot tip from Isidor Saric that he had seen one along one of the forest tracks we had walked along a couple of days earlier, but at 3PM. He was spot on with the location though, and, to our delight, it was still present, a few metres from where we started to search! A great bit of spotting by Jasmin, as it flew down from the forest canopy to a tiny ditch behind us.

Easily overlooked on the ground among the debris of the small spring-fed ditch.

The only wet areas of the dirt track were where spring water from uphill appeared at its edge, there had not been any rain recently in what was turning into a prolonged dry spell. We watched it more or less continuously from 1145 to 1303 on 15 June, taking salts, or perched up in the canopy, at around 4-5m from the ground, usually on leaves but a couple of times on a tree trunk. At 790m ASL this sighting was at the lower end of the altitudinal range and well short of the regular 1569m ASL Babin Zub Hotel site. The temperature was around 25 degrees Celsius, and it was noticeable that it only flew to the ground after basking in sunshine. When the sun was occasionally covered by a cloud it remained in the leaves. It also stuck to a stretch of around only 20m, and mostly flew up or down, rather than along the track, so it would be very easy to miss if our walk by did not coincide with one of its visits to the ground, as had presumably happened already. We had seen plenty of other butterflies taking salts along this track, including numerous Commas Polygonia c-album but had drawn a blank with False Comma until now. When taking salts, it could be approached quite closely with care, but it did not like rapid movements. A very special close encounter with one of Europe’s rarest butterflies!

False Comma, taking salts from a trackside ditch.

Its distribution is not limited by the source of food as their caterpillars have quite a wide range of food plants including common birch, willow, poplar and elm species. So, the recent decline in Europe is thought to be climate related. False Commas are thought to have quite specific climate requirements, favouring prolonged and stable sub-zero temperatures during hibernation (in such places as old wood stacks and disused buildings) but they do not do well when early warm spells are followed by severe frosts. When they emerge, they do not seek nectar but other substances including sap and animal excrement. Eggs are laid in shady and damp spots in woodland, not in direct sunlight as with some other nymphs. N. vau-album is also strongly migratory (especially to the east of Europe) and this may have influenced records in eastern Europe but the consistent presence of fresh adults in Serbia means it is certainly breeding there and therefore must be a habitat and climatic sweet spot.

The spectacular False Comma, a life-changing experience for butterfly enthusiasts!

False Comma retreats to the trees when disturbed, or just when it has taken on enough salts.

False Comma keeping an eye on us from the canopy.

 

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